Figure 11a

Lab Members



Hoddle Lab Members


Mark Hoddle

Mark S. Hoddle, Ph.D.

Biological Control Specialist and Principal Investigator 

Dr. Hoddle has headed the research in this laboratory since 1997 and is primarily involved in the identification of pest problems wf biological control could be a successful approach. The location, release and evaluation of natural enemy impacts on population growth features strongly in his research. The evaluation of biological control agents are conducted primarily in the field and, when necessary, aspects of both pest and natural enemy biology and behavior are studied in the laboratory.


Nicola A. Irvin, Ph.D.

Biological Control Specialist and Research Scholar

Dr. Irvin joined the Hoddle laboratory in 2001 as a post doctoral scholar. Dr Irvin has had a heavy focus on researching the glassy-winged sharpshooter, Homalodisca vitripennis (Germar), and its Mymarid wasp biological control agentsGonatocerus ashmeadiG. fasciatusG. triguttatus, and G. tuberculifemur. Currently,  Dr. Irvin is collaborating with Dr. Milosavljević on research investigating whether insectary plants (used to augment populations of natural enemies) and biodegradable hydrogel bead baits (for control of Argentine ants) can enhance natural enemy abundance and increase biological control of citrus pests. 

Ivan Milosavljevic

Ivan Milosavljević, Ph.D

Assistant Project Scientist

Dr. Milosavljević joined the Hoddle laboratory in 2016 as a postdoctoral researcher. His work has embraced a wide range of research projects sponsored by various granting agencies. An example of these projects include: (1) researching the Asian Citrus Psyllid (ACP), Diaphorina citri Kuwayama (Hemiptera: Psyllidae), and its biological control agents Diaphorencyrtus aligarhensis and Tamarixia radiata, (2) developing detection programs for the invasive South American palm weevil (Rhynchophorus palmarum), and (3) mitigating export risks associated with bean thrips (Caliothrips fasciatus). Dr. Milosavljević was promoted to Assistant Project Scientist in 2020. His current research focuses on the development and evaluation of novel technological tools to improve and streamline the process of controlling sap sucking citrus pests and the ants that tend them in commercial citrus.


Francesc Gomez Marco

Francesc Gomez Marco, PhD

Postdoctoral Researcher

Dr. Gómez Marco joined the Hoddle laboratory as a Postdoctoral Researcher in June 2019. His previous research experience was on biological control programs and insect ecology on citrus pests principally. He has been working on the Proactive Biological Control Program for the Spotted Lanternfly (Lycorma delicatula) which has been sponsored by the Californian Department of Agriculture. The main objective of this project is to develop a biological control program with the parasitoid of the spotted lanternfly Anastatus orientalis and to test the non-target hosts of this parasitoid species on the West coast of USA.

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